Lewontin and Kojima (1960) demonstrated that epistatic fitness interactions can produce population dynamics indistinguishable from neutral models at the level of allele frequencies alone.

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Allele frequencies(in genetics): How common different versions of a gene are in a population, expressed as percentages or proportions (for example, 30% of organisms might have version A of a gene, 70% version B).
Epistatic fitness interactions(in genetics and evolution): A situation where the effect of one gene on an organism's survival and reproduction depends on what other genes it has—meaning genes don't work in isolation but influence each other.
Lewontin and Kojima(as cited researchers in population genetics): Richard Lewontin and Shoji Kojima were biologists who conducted influential research in the 1960s on how genes behave in populations; they're referenced here because their specific 1960 study is being discussed.
Neutral models(in evolutionary theory): Scientific models that assume genetic changes happen randomly rather than because they help or hurt an organism's survival—used as a baseline for comparison.

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