Selective pressures on human brains differed dramatically (social complexity, teaching, tool use); different selection pressures produce phenotypic divergence even from similar substrates.

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Phenotypic divergence(as used in evolutionary biology): When organisms that start out similar develop noticeably different visible traits or characteristics because they face different conditions.
Selective pressures(as used in evolutionary biology and anthropology): Environmental or social conditions that make certain traits more likely to help an organism survive and reproduce, so those traits become more common over time.
Substrate(as the underlying physical foundation being discussed): The physical stuff or material that something is made of or runs on; in this case, the brain as the physical basis for mental states.
social complexity(as used to explain why evaluating freedom becomes harder): The complicated nature of society created by many interacting people, institutions, and factors that influence each other.

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