The SLOSS debate (Simberloff & Abele, 1976) demonstrated empirically that several small reserves can collectively harbor more species than a single large reserve of equal total area.
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Two ecologists (David Simberloff and Jared Abele) who studied and published research about the best ways to design nature reserves to protect biodiversity.
empirically(as used in epistemology (how we know things))
Based on actual observations and real-world evidence rather than just ideas or beliefs.