Under Millikan's framework, a structure that has permanently lost its selected causal role is no longer a fully-fledged member of its functional natural kind, regardless of positional continuity.

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Millikan(The statement is about her specific theory): Ruth Garrett Millikan is a modern philosopher who studies how language and thoughts get their meaning, especially by looking at how living things evolved and developed purposes over time.
functional natural kind(describes what kind of thing a structure is): A category of things that belong together because they all perform the same biological or natural function, like 'hearts' or 'eyes'—they're grouped by what they do, not just what they look like.
positional continuity(contrasted with whether something truly counts as a member of its kind): When something stays in the same physical location or position, even if it stops working the way it's supposed to.
selected causal role(determines whether something counts as a genuine member of its kind): The specific job or function that something was historically used for and evolved to do—like how a heart's selected causal role is to pump blood because that's what it developed to do.

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