
1929 – 2021
Richard C. Lewontin (1929–2021) was an American evolutionary biologist and geneticist at Harvard University, renowned for his contributions to population genetics and his sharp critiques of genetic determinism and adaptationism. He co-developed the concept of the 'spandrel' critique with Stephen Jay Gould and was a leading voice in debates over the gene-centric view of evolution.
Co-authored 'The Spandrels of San Marco' with Gould, critiquing pan-adaptationism
Pioneered the use of gel electrophoresis to measure genetic variation in natural populations
Demonstrated that replicator dynamics need not converge to evolutionarily stable states
Authored 'The Genetic Basis of Evolutionary Change' (1974), foundational in population genetics
Sustained critique of genetic determinism and sociobiology in works like 'Not in Our Genes'