
1935 – 2010
Leigh Van Valen (1935–2010) was an American evolutionary biologist at the University of Chicago known for sweeping, unconventional contributions to evolutionary theory and paleontology. He proposed the Red Queen hypothesis and Van Valen's Law of Extinction, and engaged seriously with the philosophy of biology, including scrutiny of the logical and cognitive foundations of natural selection.
Proposed the Red Queen hypothesis (1973): organisms must continuously evolve merely to survive against co-evolving competitors
Formulated Van Valen's Law of Extinction: extinction probability remains roughly constant over time within a clade
Founded the journal Evolutionary Theory (1974) as a venue for foundational and heterodox work
Challenged the cognitive status of natural selection, questioning whether it is genuinely predictive or empirically falsifiable
Made significant contributions to Paleocene mammalian paleontology and the classification of early placental mammals